Propublica, a non-profit online news organization, won a Pulitzer for investigative reporting for its report on New Orleans hospitals after Hurricane Katrina.
The Washington Post won awards for international reporting, commentary, criticism and feature writing. The New York Times won for national reporting and explanatory reporting.
"This is something we're going to see more of in the years ahead as there's more and more collaboration of news entities when it comes to enterprise journalism," Sig Gissler, administrator of the prizes, said in announcing the winners.
San Francisco Chronicle website SFGate.com won for editorial cartooning. The award for the animated cartoons by Mark Fiore marked the first time an Internet-based entry won in that category.
The Pulitzers are given out annually by Columbia University on the recommendation of a board of distinguished journalists and others. Each award carries a USD 10,000.